I wonder if not only has to do with intonation, but also to get a better break angle on the unwound strings at the same time.
I don't know this for a fact, but just wondering.
As far as the pickups go, Takamine's pickup system is pretty cool. They have a piston for each string that are mounted under the bridge plate, and go through it and contact the pickup in the saddle slot.
I have never played (or heard) a Takamine that was not perfectly in balance plugged in.
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